Comprehensive Description

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 Herb to 20 cm, epipetric and terrestrial. Stems procumbent and erect, drying dark brown, maroon to dark green when fresh, pubescent, more densely so towards the shoot tips, the hairs 0.5 mm, erect or weakly appressed, curved or crooked, orange-brown peltate glandular, cystoliths absent, the internodes 4–38 × 1.5–2.0 mm, angulate in cross-section, striate. Stipules 2.5–3.0 mm, auriculate-cordiform, drying brown. Leaves petiolate, distichous; petioles at each node unequal by ratio 1:3–4.4, 3–20 mm, pubescent, the hairs 0.25-0.375 mm, erect, strongly curved to curved; laminae at each node equal or subequal, 11–35 × 7–17 mm, ovate to broad ovate, chartaceous; 3-nerved, the lateral nerves visible for less than 2/3 of the lamina length, secondary nerves 4–6 pairs, borne 45–60° to the midrib, straight or weakly curved; upper surface drying brown or dark brown, dark green with maroon nerves and green flushed maroon when fresh, sparsely pubescent, the hairs 0.50–0.675 mm, appressed, straight or weakly curved, cystoliths absent, midrib raised; lower surface drying grey-brown when fresh, nerves densely pubescent, the hairs 0.375 mm, weakly appressed, curved, orange-brown peltate glandular, midrib and lateral nerves raised, cystoliths fusiform, randomly scattered; base symmetrical, cuneate or obtuse; margin serrate, the basal 1/4 entire; apex symmetrical, subcuspidate or cuneate. Inflorescences 4–10 per stem, unisexual, staminate and pistillate inflorescences synchronous, born on separate stems; bracts 0.375 mm; bracteoles 0.3–0.5 mm. Staminate inflorescences solitary, 17.5–25 mm, bearing 7–16 flowers in a loose cyme; peduncle 1/4 or less inflorescence length, 0.5 mm in diameter, pubescent, cystoliths absent; pedicels 0.8–1.0 mm, glabrous. Staminate flowers 1.5–2.0 × 1.5–1.8 mm immediately prior to anthesis, deep pink; tepals 4, imbricate, 1.75 mm, fused for their basal 1/4, ovate or elliptic, glabrous, the subapical appendage 0.375 mm, corniculate, glabrous; stamens 4. Pistillate inflorescences solitary, 2.0–2.5 mm, bearing 17–30 flowers in a compact cyme; peduncle 1/2 to 2/3 inflorescence length, 0.375 mm in diameter, glabrous, cystoliths absent; pedicels 0.25–0.375 mm, sparsely pubescent. Pistillate flowers 0.375–0.50 mm, tepals 3, unequal, glabrous, adaxial tepal 0.5 mm, oblong, the dorsal tepal appendage 0.375 mm, oblong, markedly thickened; the lateral tepals 0.25–0.375 mm, asymmetrically ovate. Infructescences not seen.


Creative Commons Attribution 3.0 (CC BY 3.0)

© Alex K. Monro, Y. G. Wei, C. J. Chen

Source: PhytoKeys

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