Phalanger lullulae (Woodlark Island Cuscus) is only found on Woodlark Island, which is part of the Milne Bay Province of Papua New Guinea, and on the neighboring island of Alcester, which is 70 kilometres south of Woodlark (Norris, 1999).

Biogeographic Regions: australian (Native ); oceanic islands (Native ); pacific ocean (Native )

Other Geographic Terms: island endemic

Phalanger lullulae has a distinctive pelage. It is short and woolly with irregularly marbled brown, ochre, and white dorsal areas. The ventral fur is white with irregular dark spots. The color varies on individuals along a continuum from predominantly dark with some lighter spots to predominantly light colored with a few small darker spots (Flannery, 1995). The species has black facial skin and a pink rhinarium. Pale ear flashes are sometimes present.

Phalanger lullulae is a medium sized marsupial. The females are on average slightly larger than the males.

Phalanger lullulae is highly adapted to arboreal life. The tail is long and prehensile. The end of the tail is naked and used to assist in gripping. Digits one and two are opposable against three, four, and five. The first and second digits of the pes are syndactylous.

The skull is pear shaped and widest at the posterior end of the zygomatic arch. With age the supraorbital ridges fuse to form a sagittal crest. The intraorbital trough is broad and shallow. The lacrimal is broadly exposed on the face of the rostrum (Norris, 1999).

The dental formula is 3/1 1/0 2/1 4/4=32. There are also two to three unicuspids of unknown homology between i1 and p3 (Menzies, 1986). The molars are not strongly crenulated. There is a well developed paraconid on m2 (Norris, 1999).

Range mass: 1500 to 2050 g.

Average mass: 1730 g.

Range length: 638 to 717 mm.

Average length: 688 mm.

Other Physical Features: endothermic ; bilateral symmetry

• Terrestrial

Phalanger lullulae prefers primary and secondary lowland dry forest. For this reason, it is more plentiful on the eastern side of Woodlark Island, where this is the predominant type of vegetation, than in the dense jungle of the western side (Flannery, 1995).

Habitat Regions: tropical

Terrestrial Biomes: forest

Local people on Woodlark and Alcester Islands claim that P. lullulae feeds on two species of vine. The species of these vines have not yet been identified. It has been suggested based on information from other species of Phalanger that they may also eat fruit and even meat when available (Flannery, 1995).

Primary Diet: herbivore (Folivore )

Phalanger lullulae may compete with the omnivorous sugar gliders and frugivorous bats (Dobsonia pannietensis, Nyctimene major, Pteropus conspicillatus, Pteropus hypomelanus- Flannery, 1995) which also forage in the forest canopy for food. It is not known to raid gardens, so it is not regarded as a pest (Norris, 1999).

No anti-predator adaptations have been described for Phalanger lullulae. Phalanger lullulae is the largest species of terrestrial mammal (with the exception of humans) on Woodlark Island, so it may have no predators in the wild (Norris, 1999). The animal is hunted by the people of the island (Flannery, 1995).

Known Predators:

• humans (Homo sapiens)

Little is known about the specific development and life cycle of P. lullulae. However, like many marsupials the young are born naked and highly altricial. The young are carried in the marsupium, after which they grasp onto the mother's back and ride there while they continue to mature (Vaughn, Ryan, and Czaplewski, 2000; Norris, 1999).

Mating behavior has not been observed in P. lullulae. The only information on reproduction and ontogeny comes from the capture of five females in August 1987. Of these females, one was parous, one had no young, two had naked pouch young, and one had a well grown back young. From this it can be inferred that the breeding season is an extended period (Flannery, 1995).

Key Reproductive Features: gonochoric/gonochoristic/dioecious (sexes separate); sexual ; viviparous

Initial expeditions which concentrated on the western side of Woodlark Island found P. lullulae to be scarce, leading to fears that the species was endangered. More recent expeditions have found it to be moderately abundant on the eastern side of the island and on Alcester Island, even around human populations where it is hunted. The species is still considered vulnerable because of it's limited geographic distribution (Norris, 1999).

US Federal List: no special status

CITES: no special status

IUCN Red List of Threatened Species: endangered

Phalanger lullulae is hunted for meat by Woodlark Island's indigenous people. However, the meat from P. lullulae is only a minor part of the local diet, and the animals are only hunted when the sea is too rough to fish (Flannery, 1994).

Positive Impacts: food