Comprehensive Description

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 ♂: Total body length 9.9 mm (7.5–9.9 mm); forewing length 7.4 mm (6.0–7.9 mm). Head wider than long (length 2.4 mm, width 3.0 mm); upper interorbital distance 1.8 mm; lower interorbital distance 1.3 mm. Intertegular distance 2.4 mm (2.0–2.6 mm); mesoscutellar posterior margin often sinuate, sometimes weakly so, such that apicolateral angle projects as a prominent, broad spine and with a defined median emargination [degree of this sinuation is variable and so some males have spines and median emargination less prominent, but margins from apicolateral corners to midpoint are never straight as is usual for species such as Thyreus hohmanni Schwarz, typical Thyreus ramosus (Lepeletier de Saint Fargeau), or several Asiatic species]. Inner anterior angle of metatibia not swollen or projecting into prominence or point between metatibial spurs (e.g., in some Palearctic species this area of metatibia is prominently developed: e.g., Lieftinck 1968). Apex of seventh metasomal tergum with apicolateral prominences distinct, truncate margin between straight, without medial emargination or swelling; male terminalia as in Figures 7–11.  Labrum with coarse, irregular, contiguous punctures except basolateral impunctate areas, such areas longer than wide and therefore ovoid in shape, basomedially with short V-shaped furrow; clypeus with small contiguous punctures, integument between (where evident) smooth; face as on clypeus except punctures slightly larger, nearly contiguous, and somewhat weaker on supraclypeal area; punctures weaker and shallower on vertex and on ocellocular area, with small impunctate area bordering lateral ocellus; punctures coarse, shallow, and nearly contiguous on gena and posterior area of postgena, anterior area of postgena smooth and impunctate. Pronotum with coarse, shallow, nearly contiguous punctures; mesoscutum with well-defined punctures separated by less than a puncture width, slightly more widely spaced around parasidal lines and medioposteriorly such that punctures are separated by about 0.5–2 times a puncture width, integument between punctures smooth and shining; mesoscutellum, including axilla, with punctures as on medioposterior section of mesoscutum except punctures separated by 0.5–1 times a puncture width; pleura with coarse, nearly contiguous punctures, integument between punctures (where evident) smooth and shining; hypoepimeral area with impunctate area bordering scrobe; propodeal lateral and posterior surfaces with coarse, shallow, ill-defined, nearly contiguous punctures. Metasoma with small punctures separated by less than a puncture width, punctures coarser, larger, and somewhat more poorly defined on more apical terga, integument between finely imbricate, apical margins narrowly impunctate and finely imbricate; sterna with similar punctation except those on discs of more basal sterna more widely spaced and becoming more poorly defined on more apical sterna.  Integument black except dark brown on tarsi, mouthparts, and apically on seventh metasomal tergum and on apical sterna. Wing membranes hyaline and slightly infumate, veins dark brown to black.  Pubescence generally fuscous to black over entire body except for presence of plumose white setae on face (Fig. 5), posterior of vertex, upper gena, outer surface of protibia, outer surface of mesotibia, outer basal surface of metatibia, and on mesosoma (using annotation system of Lieftinck 1962, 1968) as follows: deps (dorsal mesepisternal) and lpn (lateral pronotal) present; als (anterolateral mesoscutal) present but reduced, often faint; ms (median mesoscutal) present, although often reduced and faint; mls (mediolateral mesoscutal) present; plsa (anterior posterolateral mesoscutal) present along anterior half to two-thirds of border with tegula, not meeting pls (posterolateral mesoscutal) posteriorly; t (tegular) present and prominent posteriorly on tegula; pls (posterolateral mesoscutal) present, not extending laterally to meet plsa (anterior posterolateral mesoscutal); ps (parascutellar) and s (mesoscutellar) absent; deps (dorsal mespisternal), hypm (hypoepimeral area), and lp (lateral propodeal) present, veps (ventral mesepisternal) absent (Figs 1, 3). Mesoscutellum with dense patch of long, plumose, white setae extending posteriorly from undersurface of mesoscutellum medially, patch wide but not reaching to apicolateral corners. Metasomal terga with prominent patches of appressed, plumose white setae as follows: first metasomal tergum with large, L-shaped patches laterally; second metasomal tergum with lateral patch transverse, slightly wider than twice as long, never L-shaped and without rounded secondary anterior patch; third through sixth metasomal terga with more or less transverse to rounded lateral patches (Figs 1, 3).  ♀: As described for the male except in usual gender differences and as follows: Total body length 8.1–10.6 mm; forewing length 6.4–7.4 mm. Head wider than long (length 2.4 mm, width 3.0 mm); upper interorbital distance 1.9 mm; lower interorbital distance 1.3 mm (Fig. 6). Intertegular distance 2.1–2.5 mm; mesoscutellar posterior margin as in male but sometimes sinuate margin weaker and apicolateral angle not forming as prominent a spine. Apical depression of fifth tergum densely punctate and setose. Pygidial plate relatively narrow, margins converging apically, largely straight, apex narrowly rounded, surface imbricate, basal half with shallow, coarse punctures.  Clypeus with small punctures more widely spaced than in male, separated by less than a puncture width, punctures variable in size placed apically, but rather uniform medially, integument between smooth. Mesoscutum with well-defined punctures separated by less than a puncture width anteriorly and laterally, slightly more widely spaced on disc and posteriorly, there separated by 0.5–1.5 times a puncture width; mesoscutellum with punctures separated by less than a puncture width, less frequently separated by as much as a puncture width, axilla with punctures separated by less than a puncture width.  Integument and pubescence as in male except dark brown on pygidial plate; second through fifth metasomal terga with more or less transverse to rounded lateral patches (Figs 2, 4).


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© Jakub Straka, Michael S. Engel

Source: ZooKeys

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