The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 2004. database, Amphibian Species of the World: an Online Reference
  • Ardila-Robayo, M. C. (2005). ''Atelopus angelito.'' Ranas Arlequines. Rueda-Almonacid, J. V., Rodríguez-Mahecha, J. V., Lötters, S., La Marca, E., Kahn, T. R., and Angulo, A., eds., Conservación Internacional. Panamericana Formas e Impresos S. A., Bogotá, Colombia.
  • Ardila-Robayo, M. C. and Ruiz-Carranza, P. M. (1998). ''Una nueva especie de Atelopus A. M. C. Dumeril & Bibron 1841 (Amphibia: Bufonidae) de la Cordillera Central Colombiana.'' Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 22(83), 281-285.
  • Coloma, L.A., Duellman, W.E., Almendáriz, A., Ron, S.R., Terán-Valdez, A., and Guayasamin, J.M. (2010). ''Five new (extinct?) species of Atelopus (Anura: Bufonidae) from Andean Colombia, Ecuador, and Peru.'' Zootaxa, 2574, 1-54.
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • La Marca, E., Lötters, S., Puschendorf, R., Ibáñez, R., Rueda-Almonacid, J. V., Schulte, R., Marty, C., Castro, F., Manzanilla-Puppo, J., García-Pérez, J. E., Bolaños, F., Chaves, G., Pounds, J. A., Toral, E., and Young, B. E. (2005). ''Catastrophic population declines and extinctions in neotropical harlequin frogs (Bufonidae: Atelopus).'' Biotropica, 37(2), 190-201.
  • Lötters, S. (2007). ''The fate of the Harlequin Toads — help through a synchronous multi-disciplinary approach and the IUCN ‘Amphibian Conservation Action Plan’.'' Zoosystematics and Evolution, 83( Supplement 1), 69-73.
  • Merino-Viteri, A. (2001). ''Análisis de posibles causas de las disminuciones de poblaciones de anfibios en los Andes del Ecuador.'' Tesis de Licenciatura, Quito, Ecuador, Pontificia Universidad Católica del Ecuador,
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Stuart, S., Hoffmann, M., Chanson, J., Cox, N., Berridge, R., Ramani, P., and Young, B. (eds) (2008). Threatened Amphibians of the World. Lynx Edicions, IUCN, and Conservation International, Barcelona, Spain; Gland, Switzerland; and Arlington, Virginia, USA.
  • Yánez-Muñoz, M. and Altamirano B., M. (2005). ''Atelopus sp. 14.'' Ranas Arlequines. Rueda-Almonacid, J.V., Rodríguez-Mahecha, J. V., LaMarca, E., Lötters, S., Kahn, T., and Angulo, A., eds., Conservación Internacional. Panamericana Formas e Impresos S. A., Bogotá.


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