Comprehensive Description


The following pictures are of larvae of the genus Coptoclava, from the Early Cretaceous of the Shandong Province of China.


Paddle-like mesothoracic and metathoracic legs

End of abdomen


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This Mesozoic family consists of medium-sized to large aquatic beetles with divided compound eyes. Larvae have raptorial front legs. The subfamily Necronectinae is found in the Middle to Late Jurassic of Asia and Europe, and Early Cretaceous of northern Africa, Coptoclavinae in the Early Cretaceous of Asia, and Charonoscaphinae in the Late Jurassic of Asia. See Ponomarenko (1977, 1992) for further details.


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Coptoclavidae is an extinct family of beetles in the suborder Adephaga.[1][2] The Coptoclavidae lived from the Jurassic to the Early Cretaceous.[3]


  1. ^ Tree of Life Web Project. 1995. Liadytidae. Liadytes. Version 1 January 1995 (temporary). [1].01.01 in The Tree of Life Web Project, [2]
  2. ^ Ivan Löbl & Aleš Smetana (May 2003). "1". Catalogue of Palaearctic Coleoptera: Archostemata, Myxophaga, Adephaga 1. Apollo Books. pp. 6–8. ISBN 978-87-88757-73-6. 
  3. ^ Grimaldi, David and Michael S. Engel (2005-05-16). Evolution of the Insects. Cambridge University Press. ISBN 0-521-82149-5. 
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