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DescriptionPerennial herbs, creeping or climbing. Stems 4-angled, brittle. Stems, leaf margins and petioles with recurved prickles, the whole plant tending to cling to clothing. Stipules similar to the leaves. Leaves (and leaf-like stipules) in whorls of 4-8, petiolate. Flowers in axillary or terminal cymes, bisexual, usually 5-merous. Calyx reduced to a rim; teeth 0, not enclosed in an involucre. Corolla rotate to ± campanulate, greenish to yellow. Ovary 2-locular with 1 ovule per loculus. Fruit fleshy, glabrous, consisting of two round, 1-seeded mericarps.