The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 1. National Biodiversity Network Species Dictionary (January 2004) External link.
  • 2. Harde, K.W. (2000) Beetles. Silverdale Books, Leicester.
  • 3. Chinery. M. (1993) Insects of Britain and Northern Europe. Harper Collins Publishers Ltd, London.
  • 4. Kendall Bioresearch Services (January 2004) External link.
  • 5. HYPP Zoology (January 2004) External link.
  • 6. Buczaki, S. (2002) Fauna Britannica. Hamlyn, London.
  • Arnett, R. 1985. American Insects. New York: Van Nostrand Reinhold.
  • Blum, M. 1985. Fundamentals of Insect Physiology. New York: John Wiley and Sons.
  • Burton, J. 1979. The Oxford Book of Insects. New York: Oxford University Press.
  • Fraval, A. 1998. "HYPP Zoology" (On-line). Accessed April 15, 2001 at
  • Jessop, L. 1986. Dung beetles and chafers (Coleoptera: Scarabaeoidea). RES Handbk. Ident. Br. Insects, Vol. 5, Part 11.
  • Linnaeus C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I. Editio X. Laurentii Salvi, Holmiae :1-824
  • Melolontha melolontha (L., 1758)
  • Pesson, P. 1959. The World of Insects. New York: McGraw-Hill.
  • Schoolmeesters P. (2015). Scarabs: World Scarabaeidae Database (version Jul 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.


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