The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 1996. database, NODC Taxonomic Code
  • 2004. database, Amphibian Species of the World: an Online Reference
  • Amphibian Species of the World V2.20 Database. The American Museum of Natural History. Department of Herpetology.">">
  • Coloma, L.A., Ron, S., Jungfer, K-H., Grant, T., Cisneros-Heredia, D., Solís, F., Ibáñez, R., Chaves, G., Savage, J., Jaramillo, C., Fuenmayor, Q., Bolaños, F., Lips, K., Lynch, J.D., and Almendáriz, A. (2008). Gastrotheca cornuta. In: IUCN 2009. IUCN Red List of Threatened Species. Version 2009.1. Downloaded on 28 July 2009.
  • Costa Rica, Asamblea Legislativa. Decreto No. 26435-MINAE. La Gaceta, Diario Oficial (3 de Diciembre 1997).
  • Costa Rica, Asamblea Legislativa. Ley Orgánica del Ambiente, No. 7554. La Gaceta, Diario Oficial, No. 215 (13 de Noviembre 1995).
  • Costa Rica, Ministerio de Ambiente y Energía (MINAE). 1997. Ley de Conservación de la Vida Silvestre y su Reglamento, No. 7317. 2 da. Edición. EDITEC EDITORES, S.A.. San José, Costa Rica. 134 p.
  • Duellman, W.E. 1983. A new species of marsupial frog (Hylidae: Gastrotheca) from Colombia and Ecuador. Copeia 1983(4): 868-874.
  • Duellman, W.E. 1983. Gastrotheca aureomaculata: systematic resolution of a case of intrapopulational polycromatism in a hylid frog. Herpetologica 39: 105-110.
  • Duellman, W.E. 1989. Lista anotada y clave de los sapos marsupiales (Anura: Hylidae: Gastrotheca) de Colombia. Caldasia 16(76): 105-111.
  • Duellman, W.F. 1970. The Hylid Frogs of Middle America. Vol. l. Monogr. Mus. Nat. Hist. Univ. Kansas No. 1.
  • Frost, D.R. 1985. AMPHIBIANS SPECIES OF THE WORLD: A Taxonomic and Geographical Reference. Published as a Joint Venture of Allen Press, Inc. and The Association Systematics Collections Lawrence, Kansas, U.S.A. 732 p..
  • Frost, Darrel R. 1997. Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
  • Frost, Darrel R. 1997. Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Holotype: Stejneger, L. 1911. Proc. U. S. Nat. Mus. 41 (1857): 286.
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Savage, J. M. (2002). The Amphibians and Reptiles of Costa Rica. University of Chicago Press, Chicago and London.
  • Savage, J.M. 2002. The amphibians and reptiles of Costa Rica: A Herpetofauna between two continents, between two seas. The University of Chicago Press.
  • Solórzano, A., W. E. Lamar y L. Porras. 1997. The marsupial frog Gastrotheca cornuta (Amphibia: Hylidae) in Isthmian Central America. Rev. Biol. Trop. 45(4): 1691-1694.
  • Universidad de Costa Rica. s.f.. CATALOGO DE HERPETOLOGIA. Museo de Zoología, Escuela de Biología, Universidad de Costa Rica. s.d..
  • Wassersug, R. J., and Duellman, W. E. (1984). ''Oral structures and their development in egg-brooding hylid frog embryos: evolutionary and ecological implications.'' Journal of Morphology , 182, 1-37.
  • del Pino, E. (1980). ''Morphology of the pouch and incubatory integument in marsupial frogs (Hylidae).'' Copeia, 1980, 10-17.
  • del Pino, E. M. and Escobar, B. (1981). ''Embryonic stages of Gastrotheca riobambae (Fowler) during maternal incubation and comparison with development of other marsupial frogs.'' Journal of Morphology, 167, 277-295.
  • del Pino, E., and Humphries, A. A., Jr. (1978). ''Multiple nuclei during early oogenesis in Flectonotus pygmaeus and other marsupial frogs.'' Biological Bulletin, 154, 198-212.


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