Macrohabitat: Lowlands to midlands, sea level- 900 meters altitude. In Europe, this species is a pronounced forest beetle except in the humid climate of the west where it occurs in open country. Microhabitat: Clearings, roadsides, vacant lots, open grassy cultivated fields, and forest edges. Adults occur on shaded or open ground that is consisting of moderately dry gravelly, sandy, or loamy ground covered with sparse vegetation, or leaf litter. DIspersal abilities: Wing-polymorphic. European records indicate macropterous adults fly. Seasonal occurrence: Adults have been found in May-November; teneral adults in June-September. Behavior: Adults are mostly diurnal, taking cover on cloudy days under dead leaves or stones, among conifer needles or pieces of wood; they are carnivorous and in Europe, adults have been observed eating Collembola, mites, flies, and other arthropods. Larvae are known to climb on tree trunks. This species is favored by human activities and is an effective colonist. Adults hibernate. (Erwin, 2001; Larochelle & Lariviere, 2003; Lindroth, 1961; Data from CAS, UASM collections)