Overview

Distribution

Guangdong, Guangxi, Guizhou, Hubei, Sichuan, Yunnan.
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Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Sichuan, Xizang, Yunnan [N Vietnam].
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Physical Description

Morphology

Description

Shrubs 1-3 m tall. Young branchlets longitudinally angular and sulcate, yellow pubescent, glabrescent. Leaves alternate, usually congested at apices of branchlets; petiole 5-10 mm, sulcate adaxially; leaf blade green, oblong or oblanceolate, rarely ovate-lanceolate, 3-12 × 1-3 cm, subleathery, both surfaces glabrous, midvein raised abaxially, impressed adaxially, lateral veins 7-12 pairs, raised abaxially, obscure adaxially, anastomosing near margin, reticulate veins indistinct, base attenuate to cuneate, margin entire, slightly recurved, undulate, apex caudate-acuminate or long acuminate. Racemes terminal or axillary at apices of branchlets, several together corymbose or paniculate, 2.4-5(-7) cm, connate pubescent. Pedicel 1-1.5 mm, glabrous; basal bracteoles 3, caducous, deltoid-ovate. Flowers small, 5(-8) mm. Sepals 5, unequal, ciliate, caducous at anthesis; outer sepals 3, ovate, ca. 2 × 1.5 mm, outside pubescent, apex rounded; inner sepals 2, petaloid, obovate to obliquely obovate, large, 4.5(-6) × ca. 3 mm, 3-veined, base attenuate, apex retuse-rounded. Petals 3, connate in lower 3/4, white, yellow, or purple; keel longer than lateral petals, ca. 5 mm, apex with peltate appendages. Stamens 8; filaments ca. 4 mm, lower 3/4 united, forming an open staminal sheath; anthers ovoid. Disk annular. Ovary obovoid, ca. 0.8 mm in diam.; style thickening from base to apex, apex curved and 2-lobed; stigma in lower lobe. Capsule oblong-obovoid, ca. 8 mm, ca. 4 mm in diam., base attenuate, with annular disk, margin narrowly winged, apex retuse. Seeds brown-black, ovoid, ca. 1.5 mm, ca. 1 mm in diam., densely red-brown villous, hiliferous end brown-black convex. Fl. Nov, fr. May-Dec.
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Description

Shrubs or small trees, 1-4 m tall. Branchlets terete, longitudinally angular and sulcate, glandular-pubescent when young. Leaves congested at apices of branchlets; petiole 6-10 mm, adaxially sulcate; leaf blade elliptic, elliptic-lanceolate, or oblanceolate, 4-10 × 1.5-3 cm, subleathery, both surfaces glabrous, midvein raised abaxially, impressed adaxially, lateral veins 8 or 9 pairs, slightly raised abaxially, evident adaxially, anastomosing near margin, reticulate veins absent, base attenuate to cuneate, margin entire, undulate, apex acuminate to caudate. Racemes 2-5, several axillary at apices of branchlets, 3-7 cm, shortly white glandular-hairy. Flowers sparse, 1.2-2 cm. Pedicel ca. 6 mm; basal bracteoles 3, caducous. Sepals 5, unequal, caducous at anthesis; outer sepals 3, ovate, 2-3 × 1.5-2 mm, apex obtuse, ciliate; inner sepals 2, petaloid, obovate, ca. 1.3 cm, base attenuate, ciliate or not, apex rounded, concave. Petals 3, connate in lower 3/4, yellow, rarely white or purple-red; lateral petals shorter than keel, apex truncate; keel ca. 1.5 cm, apex with 2 cucullate appendages. Stamens 8; filaments ca. 1.5 cm, lower 3/4 united, forming an open staminal sheath adnate with petals; anthers narrowly ovoid. Ovary obovoid, 1.5-2 mm, ca. 1.5 mm in diam.; style ca. 1.2 cm, apically thickening and curved, apex shallowly 2-lobed; stigma in lower lobe. Capsule obovoid or cuneate, 1-1.4 cm, ca. 6 mm in diam., base attenuate, margin winged, apex retuse, mucronate, wing with transverse veins. Seeds brown-black, ovoid, ca. 2 mm, ca. 1.5 mm in diam., brown or white villous, hairs ca. 7 mm; strophiole absent. Fl. Apr-Jun, fr. May-Jul.
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Diagnostic Description

Synonym

Heterosamara caudata (Rehder & E. H. Wilson) Paiva & P. Silveira; Polygala comesperma Chodat; P. wattersii Hance (1882), not Hance (1881).
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Synonym

Heterosamara wattersii (Hance) Paiva & P. Silveira; Polygala mariesii Hemsley.
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Type Information

Isotype for Polygala caudata Rehder & E.H. Wilson in Sarg.
Catalog Number: US 458149
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Original publication and alleged type specimen examined
Preparation: Pressed specimen
Collector(s): A. Henry
Locality: Mengtze., Yunnan, China, Asia-Temperate
Elevation (m): 1600 to 1800
  • Isotype: Rehder, A. & Wilson, E. H. 1914. Pl. Wilson. 2: 161.
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Ecology

Habitat

● Forests on limestone mountains; 1000-1800(-2100) m.
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Broad-leaved forests, shrub forests on limestone mountains; 1000-1500 m.
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