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 Thallus: squamulose, 1-2 cm diam., rosette-forming, lobate; lobes: initially rounded and scattered, becoming densely aggregated and imbricate, 0.1-0.5 mm wide (rarely elongating to 2 mm); upper surface: brown when dry, pale brown to deep green when wet, smooth; isidia: marginal to submarginal, globose to cylindrical; upper cortex: paraplectenchymatous, 30-65 micrometer thick; medulla: whitish, loosely interwoven hyphae around Nostoc colonies; lower surface: brownish, rhizinate; rhizines: blackish, densely tufted, at base of the lobes; Apothecia: rather rare, brown or black, laminal or marginal, up to 1.5 mm diam., sessile to substipitate; margin: pale brown; disc: flat, brown to black, smooth; true exciple: paraplectenchymatous, 85-170 micrometer wide, outer part brown pigmented, not supported by pseudoexciple; hymenium: brown above, K-, colorless below, I+ blue, 100-175 micrometer high; paraphyses: septate, branched, c. 2 micrometer wide, ± swollen at the apices and pigmented; subhymenium: colorless to brown, 115-190 micrometer thick; asci: cylindrical, 58-90 x 8-20 micrometer, 8-spored; ascospores: fusiform to narrowly ellipsoid, hyaline, (1-) 3-septate, 17-28 x 6.5-9 micrometer; Pycnidia: brown to black above, pale below, immersed, up to 0.2 mm diam.; conidia: bacilliform or slightly bifusiform, colorless, simple, 2.5-4 x 1 micrometer; Spot tests: all negative; Secondary metabolites: none detected.; Substrate and ecology: on sandstone in moist habitats from near sea level to 800 m; World distribution: western North America from southern California to British Columbia; Sonoran distribution: rare in southern California in the Santa Monica Mountains and on Catalina Island. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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