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 Thallus: foliose to caespitose-subfruticose, irregular, very loosely adnate, up to 7 cm in diam., ± dichotomously lobate; lobes: linear, elongate, up to 2 mm broad, but usually around 1 mm, 10-20 mm long, ± plane, sometimes prostrate but more often ascending, sometimes minutely bullate, ciliate; cilia: marginal, simple, black tipped, up to 7 mm long; upper surface: gray to dark gray, usually with black spots from the abundant pycnidia; soredia and isidia absent; upper cortex: prosoplectenchymatous; medulla: white, lax, usually well developed; lower cortex: lacking throughout; lower surface: white, without a cortex, uneven, arachnoid, erhizinate; Apothecia: ± abundant, very variable in size, short stipitate, up to 5 mm diam., without cilia on the margin; disc: black, plane, often pruinose; asci: subclavate to clavate, 8-spored; ascospores: brown, l-septate, ellipsoid, Physcia-type, (16-) 18-20.5 (-21.5) x (7.5-) 8-9.5 (-11) µm; Pycnidia: ± abundant, inserted in thallus with black openings on the lobes; conidia: bacilliform, 3.5-5 x 1µm; Spot tests: cortex K+ yellow C-, KC-, P+ yellow; medulla K+ yellow, C-, KC-, P-; Secondary metabolites: cortex with atranorin and chloroatranorin; medulla with atranorin and zeorin.; Substrate and ecology: growing on tree trunks, shrubs and cacti in open situations; World and Sonoran distribution: known from ± coastal parts of Baja California, Baja California Sur and southern California.; Notes: The ± flat lobes with marginal cilia and abundant apothecia makes this common, coastal species easy to recognize. 

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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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