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 Thallus: adnate, foliose, 4-16 cm in diam., lobate; lobes: sublinear, contiguous to imbricate, elongate, plane, separate, 1-4 mm wide, rather brittle, apices truncate; upper surface: gray to yellowish gray, smooth to foveolate, shiny or dull, sometimes white pruinose, becoming cracked along prominent, effigurate pseudocyphellae; soredia: coarse to subisidiose, abundant, yellowish, commonly marginal; isidia and pustulae absent; medulla: white with continuous algal layer; lower surface: black centrally, brown peripherally, rhizinate, rhizines moderately dense, black, simple to furcate; Apothecia: rare, laminal, 2-4 mm wide; margin: usually sorediate; disc: dark brown; asci: clavate, 8-spored; ascospores: ellipsoid, 10-12 x 5-6 µm; Pycnidia: not seen; Spot tests: upper cortex K+ yellow, C-, KC- (or + orange if usnic acid in high concentration), P+ orange; medulla K+ yellow turning deep red, C-, KC-, P+ yellow; Secondary metabolites: upper cortex with atranorin and chloroatranorin and accessory usnic acid (in the soralia); medulla with salazinic acid (major), consalazinic and protolichesterinic acids (both minor).; Substrate and ecology: frequent on exposed rocks; World distribution: boreal to arctic regions in North American and Europe; Sonoran distribution: only known from one scree slope in the upper part of the Chiricahua Mountains, SE Arizona. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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