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Sexually mature adults filiform, length to 10 mm; males smaller than females. Proboscis chambers aligned in same plane; anterior chamber to 77 μm long by 46 μm
wide; posterior chamber to 123 μm long by 16 μm; armature small; stylet to approximately 5 μm long, proximal end enlarged where attached to basis; basis ovoid, to
20 μm long by 7 μm wide. Submuscular glands open to body surface by large number of separate ducts throughout body. Mucus sheath present, non-ornamented, with distinctive spiral or corkscrew shape. Intestine with deeply lobed diverticula. Sexes separate; ovaries regularly distributed as a single row on each side, between intestinal diverticula, ovarian pores indistinct; testes abundant, irregularly scattered throughout body posterior to brain/stomach; Takakura’s duct present, diameter to 19 μm. Newly hatched planuliform larvae round, 50 μm in diameter.