The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Brooks, S. 2003. Dragonflies. London: Smithsonian Books.
  • Coin, P. 2004. "Species Aeshna umbrosa - Shadow Darner" (On-line). Bug Guide. Accessed August 03, 2012 at
  • Corbet, P. 1999. Dragonflies: Behavior and ecology of Odonata. Ithaca: Comstock Publishing Associates.
  • Dunkle, S. 2000. Dragonflies through binoculars: A field guide to dragonflies of North America. Oxford: Oxford University Press.
  • Frye, M., R. Olberg. 1995. Visual receptive field properties of feature detecting neurons in the dragonfly. Journal of Comparative Physiology A: Neuroethology, Sensory, Neural and Behavioral Physiology, 177: 569-576.
  • Garrison, Rosser W. / Poole, Robert W., and Patricia Gentili, eds. 1997. Odonata. Nomina Insecta Nearctica: A Check List of the Insects of North America, vol. 4: Non-Holometabolous Orders. 551-580
  • Iowa Odonata survey, 2005. "Shadow darner - Aeshna umbrosa" (On-line). Iowa Odonata Survey. Accessed August 03, 2012 at
  • Johnson, J. 2011. "Sorting paddle-tailed and shadow darners out-of-hand" (On-line). Northwest Dragonflier. Accessed August 02, 2012 at
  • Kraus, J. 2010. Diet shift of lentic dragonfly larvae in response to reduced terrestrial prey subsidies. Journal of the North American Benthological Society, 29/2: 602-613.
  • Lung, M., S. Sommer. 2001. "Aeshnidae: The darners" (On-line). Digital Atlas of Idaho. Accessed February 02, 2012 at
  • Manolis, T. 2003. Dragonflies and damselflies of California. Berkeley: University of California Press.
  • Mead, K. 2003. Dragonflies of the North Woods. Duluth, MN: Kollath-Stensaas Publishing.
  • Montana Natural Heritage Program, 2011. "Shadow darner - Aeshna umbrosa" (On-line). Montana field guide. Accessed August 02, 2012 at
  • Needham, J., M. Westfall. 1955. A manual of the dragonflies of North America (Anisoptera): Including the Greater Antilles and the provinces of the Mexican Border. Berkeley: University of California Press.
  • Northern Prairie Wildlife Research Center, 2006. "Dragonflies and damselflies (Odonata) of the United States" (On-line). Accessed August 02, 2012 at
  • Novelo-Gutierrez, R., K. Tennessen. 2010. Description of the larva of Aeshna persephone Donnelly, 1961 (Odonata: Aeshnidae). Zootaxa, 2484: 61-67.
  • Paulson, D. 2009. "Aeshna umbrosa" (On-line). The IUCN Red List of Threatened Species(TM). Accessed August 02, 2012 at
  • Paulson, Dennis R., and Sidney W. Dunkle. 2002. A Checklist of North American Odonata including English name, etymology, type locality, and distribution. Slater Museum of Natural History, University of Puget Sound, Occasional Paper no. 56. 88
  • Pelegrin, A. 2009. "Aeshna umbrosa Walker, 1908" (On-line). America Dragonfly. Accessed August 02, 2012 at
  • Platt, A., S. Harrison. 1995. Robber fly and trout predation on adult dragonflies (Anisoptera: Aeshnidae) and first records of Aeshna umbrosa from Wyoming. Entomological News, 106/5: 229-236.
  • Rudolf, V., J. Armstrong. 2008. Emergent impacts of cannibalism and size refuges in prey on intraguild predation systems. Oecologia, 157: 675-686.
  • The Brown Family Environmental Center at Kenyon College, 2003. "Odonata: Life cycle" (On-line). Accessed August 03, 2012 at
  • Tol J. van (2015). Odonata: Global Species Database of Odonata (version Dec 2011). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Walker, E. 1953. The Odonata of Canada and Alaska. Toronto: University of Toronto Press.
  • Walker, E. M. 1908. A key to the North American species of Aeshna found north of Mexico. [I]. The Canadian Entomologist: 40 (11): 377-391, incl. pl. 10-10.


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