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  Vine, trailing along ground or climbing on other vegetation to 3 m or more, rooting at the nodes. Stems slender, woody, moderately to densely covered with crisped transparent to tawny pubescence of unbranched, eglandular, multicellular trichomes. Sympodial units plurifoliate, not geminate. Leaves simple to 7-pinnate, most commonly 3–5-pinnate, the blades 0.8–12 × 0.5–8 cm, chartaceous, moderately to densely pubescent adaxially and abaxially, sand punctate abaxially, the rachis densely pubescent, the margins entire to irregularly revolute resulting in somewhat undulate margins, the leaflets decreasing markedly in size toward the base of the leaf, the distal leaflet of the lowermost pair typically smaller than its match or completely absent; interjected leaflets absent; lateral leaflets 0.3–3.5 × 0.2–1.5 cm, ovate to elliptic, the bases rounded to truncate, oblique, the apices obtuse to acute, the petiolules nearly lacking to 2 mm long, moderately to densely pubescent; apical leaflet 1.2–8(–9.5) × 0.7–3(–4) cm, ovate to elliptic to oblong, the apex obtuse to acute to acuminate, the base obtuse to truncate to cordate, the petiolules nearly lacking to 7 mm long, moderately to densely pubescent; petioles 0.1–2.5(–4.5) cm, moderately to densely pubescent. Pseudostipules present at most nodes, one per node, 0.5–1.5 × 0.4–0.8 cm, obliquely ovate to elliptic, sometimes lunate, the apices obtuse to acute, the bases sometimes strongly lobed, oblique. Inflorescence 1.5–3.5 × 1–3 cm, extra-axillary on main stems or terminal on short, axillary spur shoots, simple to sometimes once branched in the extra-axillary inflorescences, with 1–8 flowers (1–3 on spur shoots [mean= 1.9], 3–8 on main stems [mean = 4.6]), with all flowers apparently perfect, the axes densely pubescent; peduncle 0.5–1 cm long; rachis nearly lacking to 1.5 cm; pedicels 7–15 mm in flower, 10–20 mm in fruit, somewhat expanded distally in flower and fruit, spaced contiguously to 6 mm apart, articulated at the base. Spur shoots 0.5–3.5(–8) cm long, bracteate, with 2–8 bracts per shoot, the bracts similar in shape to the cauline leaves, simple to occasionally 3–5-pinnate, 1–15 mm long, with minute pseudostipules. Calyx 4–5 mm long, the tube 1–2.5 mm long, the lobes 2.5–3.5 × ca. 1.5 mm, ovate-lanceolate to oblong, acute at tips, moderately pubescent, sand punctate; fruiting calyx slightly accrescent, the lobes 4–4.5 × 1.5–2 mm, ovate-lanceolate to oblong. Corolla 0.8–1.5 cm in diameter, 2–6 mm long, pentagonal, white to violet, sometimes with yellow at tips or along the midveins of lobes, flat to strongly reflexed at anthesis, the lobes 1.5–5 × 3–5 mm, acute at apices, glabrous adaxially, moderately to densely pubescent abaxially along midvein of lobes, the trichomes becoming shorter toward the densely pubescent apices of the corolla lobes, the margins densely ciliate apically. Stamens equal, with filaments 0.5–1.5 mm long, nearly free to fused for about ½ their lengths, somewhat broadly flattened, nearly glabrous abaxially, densely pubescent adaxially and on margins; anthers 3–4.5 × 1–1.2 mm, oblong, incurved, connivent, yellow, the pores large, directed distally, opening into latrorse-introrse longitudinal slits with age. Ovary glabrous to sparsely pubescent; style 5–7 × 0.1–0.2 mm, exceeding stamens by 1.5–5 mm, cylindrical, glabrous to papillose in lower ½ to sparsely pubescent with long trichomes in the middle or in the lower ½; stigma capitate. Fruits 2–2.5 × 1.5–2 cm, ellipsoidal, rounded to very slightly obtusely pointed at apex, green with darker mottled striping when immature, orange when mature, glabrous to sparsely pubescent when young. Seeds 3–4.2 × 2–4 mm, flattened, lenticular, rounded to teardrop-shaped, with a 0.2–1 mm wide wing around the margins, the thickened part of the seed 1.8–2.2 × 1.5–2 mm, rounded to reniform, light to medium brown, the surface smooth, the wing yellowish-tan to transparent near the margins, with radial striations.

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© Eric. J. Tepe, Glynis Ridley, Lynn Bohs

Source: PhytoKeys

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