IUCN threat status:

Not evaluated

Comprehensive Description

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Description

  Shrub, 0.5–1.5 m. Young stems erect, slender, moderately stellate-pubescent to glabrescent, with porrect sessile or variously stalked trichomes, the stalks to 0.2 mm long, the rays 6–8, ca. 0.2 mm long, the midpoints approximately the same length as the rays, armed with straight prickles 3–4 mm long, 1–2 mm wide at base, deltate, flattened, pale yellow-orange, glabrous, spaced 5–20 mm apart; bark of older stems glabrescent, green-brown to dark brown. Sympodial units plurifoliate. Leaves lobed; blades 8–20 cm long, 5–15 cm wide, 1.5–2 times longer than wide, elliptic, chartaceous, drying concolorous to weakly discolorous, green-brown, moderately stellate-pubescent on both surfaces, with porrect, sessile or stalked trichomes, the stalks to 0.2 mm long, the rays 6–8, 0.2–0.5(-0.8) mm long, the midpoints approximately the same length as the rays, with 5–20 prickles on both surfaces; the primary veins 4–6 pairs, the tertiary venation clearly visible abaxially and not visible adaxially; base cuneate to truncate; margins lobed, the lobes 3–4 on each side, 1–3 cm long, deltate, apically obtuse to acute, extending approximately 1/3 of the distance to the midvein, often with secondary lobing; apex obtuse to acute; petiole 1–3 cm long, approximately 1/6 of the leaf blade length, moderately stellate-pubescent, with 0–5 prickles. Inflorescences apparently lateral, 3.5–9 cm long, rarely branched, with 6–15(-20) flowers, 1–4 flowers open at any one time, weakly stellate-pubescent, with 0(-5) prickles; peduncle 1–3 mm long; pedicels 1–2.3 cm long in long-styled flowers, 0.8–1.2 cm long in short-styled flowers, erect to pendent, articulated at the base, moderately stellate-pubescent to glabrescent, with 0–20 prickles on long-styled flowers, unarmed on short-styled flowers; pedicel scars spaced 2–8 mm apart. Flowers 5-merous, heterostylous and the plants andromonoecious, with the lowermost 1–3 flowers long-styled and hermaphroditic, the distal flowers short-styled and functionally male. Calyx 11–22 mm long in long-styled flowers, 5–9 mm long in short-styled flowers, the lobes 7–10 mm long in long-styled flowers, 3–4 mm long in short-styled flowers, ovate and foliaceous in long-styled flowers, deltate in short-styled flowers, apically bluntly acute in long-styled flowers and acute to obtuse in short-styled flowers, moderately stellate-pubescent, with 30–80 prickles in long-styled flowers and 0–30 prickles in short-styled flowers. Corolla 2.5–3.3 cm in diameter in long-styled flowers, 1.5–2.5 cm in diameter in short-styled flowers, usually white or white with purple midveins, sometimes mauve, stellate, lobed for 1/4–1/2 of its length, the lobes ca. 7 mm long, ca. 10 mm wide in long-styled flowers, 6–10 mm long and 5–8 mm wide in short-styled flowers, broad-deltate, spreading, sparsely stellate-pubescent abaxially, the trichomes porrect, sessile or stalked, the stalks to 0.2 mm, the rays 5–8, 0.2–0.4 mm long, the midpoints approximately the same length as the rays. Stamens equal, with the filament tube 1–3 mm long, the free portion of the filaments ca. 0.5 mm long; anthers 5–6 mm long in long-styled flowers, 4.5–5.8 mm long in short-styled flowers, connivent, tapering, poricidal at the tips. Ovary glabrous, with a few stellate trichomes towards the apex; style 1.1–1.2 cm long in long-styled flowers, stout, straight or gently curved, moderately stellate-pubescent for most of its length. Fruit a spherical berry, 1(-2) per infructescence, 2.7–3.5 cm in diameter, the pericarp smooth, dark green with pale green and cream markings when young, yellow at maturity; fruiting pedicels 2–3 cm long, 1.2–2.2 mm in diameter at base, woody, pendulous, with 0–20 prickles; fruiting calyx not accrescent, covering 1/4–1/3 of the mature fruit, reflexed, with 10–80 prickles. Seeds ca. 100–200 per berry, 2.7–3.5 mm long, 2–2.5 mm wide, flattened-reniform, orange-brown.

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Creative Commons Attribution 3.0 (CC BY 3.0)

© M.S. Vorontsova, S. Knapp

Source: PhytoKeys

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