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A monotypic tribe from the highlands of Peru. For many years, Styx was considered to be a "missing link" among butterflies, perhaps shedding light on the relationship among riodinids, lycaenids and nymphalids. It has now been shown based on DNA evidence (Wahlberg et al. 2005) to be a riodinid.Adults are pale grayish white with brown wing veins, resembling small Parnassius butterflies (Papilionidae).