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The two main apomorphies of the group are the fusion of aedeagus, juxta and anellus into a single structure, and the presence of a single ring of scales on the flagellomers. The male genitalia shows a tendency to reduce dorsal components in favor of more developed ventral structures. So uncus and tegumen are comparatively reduced compare to Tortricinae and Chlidanotinae. A part from that some venation characters are relatively primitive as the presence of chorda and M stem on forewing.


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Source: LepTree

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