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Traditional "Limenitidinae" of authors (e. g., Harvery 1991) was a paraphyletic group composed of multiple clades: Biblidini, Cyrestini, Pseudergolini, Coloburini, and Limenitidini. The latter tribe, which is the sister group of Heliconiinae, is here considered to be the "true" Limenitidinae. The rest are more closeley-related to Apaturinae and Nymphalinae (or part of Nymphalinae, in the case of Coloburini).


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Source: LepTree

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