Wikipedia
Scarabaeinae
The scarab beetle subfamily Scarabaeinae consists of species collectively called true dung beetles. Most of the beetles of this subfamily feed exclusively on dung. Well-known members include the genera Scarabaeus and Sisyphus, and Phanaeus vindex.
Scarabaeinae tribes, with some selected genera and species, are:
- Canthonini
- Circellium Latreille, 1825
- Dicranocara Frolov and Scholtz, 2003
- Versicorpus Deschodt Davis and Scholtz, 2011
- Deltochilum
- Coprini
- Copris Geoffroy, 1762
- Heliocopris
- Dichotomiini
- Eucraniini
- Eurysterini
- Gymnopleurini (sometimes included in Scarabaeini)
- Gymnopleurus Illiger, 1803
- Oniticellini
- Euoniticellus Janssens, 1953
- Oniticellus Serville, 1825
- Onitini
- Bubas Mulsant, 1842
- Cheironitis Lansberge, 1875
- Onitis Fabricius, 1798
- Onthophagini
- Phanaeini MacLeay, 1819
- Scarabaeini
- Sisyphini (sometimes included in Scarabaeini)
Coprini, Oniticellini, Onitini, Onthophagini and Phanaeini are sometimes separated as subfamily Coprinae.
Copris elphenor female
Copris elphenor male
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