Molecular Biology and Genetics

Molecular Biology

Statistics of barcoding coverage

Barcode of Life Data Systems (BOLD) Stats
Specimen Records:2091
Specimens with Sequences:1901
Specimens with Barcodes:1822
Species With Barcodes:183
Public Records:1561
Public Species:173
Public BINs:308
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)


Article rating from 0 people

Default rating: 2.5 of 5

Barcode data

Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)


Article rating from 0 people

Default rating: 2.5 of 5



The Helicidae, sometimes known as the typical snails, are a taxonomic family of small to large, air-breathing, land snails. In other words, they are terrestrial pulmonate gastropod mollusks.

A number of species in this family are valued as food items, including Cornu aspersum (formerly Helix aspersa) the garden snail, and Helix pomatia the "escargot".[1] The biology of these two species in particular has been much studied.

Shell description[edit]

The shells of these snails are mostly rather globular in shape. In some genera, such as Cepaea, the shells are brightly colored and patterned.


The animal is capable of complete retraction within the shell. The tail is without a mucous gland or projection.[2][unreliable source?]

The mouth is always provided with a jaw, which is striate, ribbed, sulcate or plicate, sometimes composed of several imbricating pieces. The radula is composed of many transverse horizontal series of teeth, the centrals tricuspidate, about the size of the laterals, laterals bicuspid, or tricuspid with the interior cusp obsolete, marginals usually wider than high, short with two or three small cusps.[2][unreliable source?]

In the soft parts the most obvious distinction is the lack of a caudal mucous pit, and their possessing a sculptured jaw. Typically, there is a distinction in the dentition, although in some species the lateral teeth take on a pseudozonitoid appearance; even in such cases the extreme marginals in Helix remain short and very obtuse.[2][unreliable source?]

This family of snails is defined by the anatomical presence of a diverticulum. Some genera within this family create and use love darts as part of their mating behavior. These snails have one dart apparatus. The dart sac has no accessory sac, and contains two tubular glands, inserted at the base of the dart sac.

In this family, the number of haploid chromosomes lies between 21 and 30 (according to the values in this table).[3]

Helicid snails for sale as food in Italy; on the left Cepaea hortensis.


The family is native to Eurasia and belongs the Western Palearctic group.

However some species, notably Cornu aspersum, have been introduced and become established in numerous different areas worldwide.

Many accidental introductions happen because the eggs of most Helicidae are laid in the soil, and they may thus easily travel unnoticed along with landscaping plants.


The family Helicidae contains 2 subfamilies (according taxonomy of the Gastropoda by Bouchet & Rocroi, 2005) and genera within the family Helicidae include (after Fauna Europaea[4] and Taxonomy Browser[5]):

Subfamily Helicinae Rafinesque, 1815 - In this subfamily the glands are divided. The love dart has four blades or vanes. There are two penial papillae.

  • Tribe Thebini Wenz, 1923 - The synonym Euparyphinae Perrot, 1939 has been declared an invalid name.
Theba Risso, 1826
Cantareus Risso, 1826
Cepaea Held, 1838
Cornu Born, 1778
Eobania P. Hesse, 1913
Helix Linnaeus, 1758 - type genus
Idiomela T. Cockerell, 1921
Otala Schumacher, 1817
Marmorana W. Hartmann, 1844 - This genus might actually belong to the subfamily Ariantinae and not to the subfamily Helicinae, as has been assumed until now.[6]

Subfamily Ariantinae Mörch, 1864 - in this subfamily the glands are divided or undivided. The love darts contain two to four blades. There is only one penial papilla.

Arianta Turton, 1831
Chilostoma Fitzinger, 1833
Cylindrus Fitzinger, 1833
Helicigona A. Férussac, 1821
Isognomostoma Fitzinger, 1833


  1. ^ M.P. Kerney & R.A.D. Cameron. 1979. A field guide to the land snails of Britain and northwestern Europe. Collins, London.
  2. ^ a b c Tryon G. W. 1887 Manual of conchology; structural and systematic. With illustrations of the species. Second series: Pulmonata. Volume 3. Helicidae - Volume I. page 3-4.
  3. ^ Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
  4. ^ Fauna Europaea
  5. ^ Taxonomy Browser
  6. ^ Nordsieck H. (2006). Higher classification of Helicoidea and the molecular analyses of their phylogeny., Accessed 20/06/12
Creative Commons Attribution Share Alike 3.0 (CC BY-SA 3.0)

Source: Wikipedia


Article rating from 0 people

Default rating: 2.5 of 5


EOL content is automatically assembled from many different content providers. As a result, from time to time you may find pages on EOL that are confusing.

To request an improvement, please leave a comment on the page. Thank you!