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 Life habit: lichenized; Thallus: crustose, continuous or slightly rimose; surface: variable, pale yellow, ochre or pale brown, smooth, waxy, rarely white or cream-colored, without soralia or isidia; cortex: mostly well developed and consisting of whitish to yellowish periclinal hyphae; medulla: sometimes with large crystals; photobiont: primary one a Trentepohlia green alga, secondary one absent; Ascomata: apothecial, lirellate or ±rounded, in some species embedded in carbonized stromata with white or pale brown surface, ascocarps or stromata usually distinctly raised; disc: with conspicuous brownish granular surface, widely open (only in G. substriatula narrow and slit-like); margin: usually present but often inconspicuous, excipular lips black, entire (crenate only in Glyphis substriatula); exciple: completely carbonized, the excipular base may be poorly developed or not carbonized (in species belonging to subgenus Pallidoglyphis), excipular lips (labia) usually entire (slightly crenate only in Glyphis substriatula), outside often with crystals embedded; hymenium: not inspersed, gelatinous, often I+ pale violet or grayish blue; paraphyses: parallel, the outer wall layers swollen, often anastomosing, tips brown-granular; asci: clavate, I-, apex thickened with ±distinct ocular chamber ('Graphis-type'), 2-8-spored; ascospores: hyaline (old or ±degenerated spores may be pale brown as well), transversely septate or muriform, oblong or oval, often with ±distinct halo, I+ blue-violet; Conidiomata and conidia: not seen; Secondary metabolites: none detected; Geography: tropical to subtropical; Substrate: bark, very seldom on leaves (only; Notes: Glyphis is characterized by a carbonized exciple, brownish granular discs, hyaline ascospores and a special paraphyses-type (parallel but often anastomosing with ±swollen outer wall layers and brown-granular tips). Glyphis is a synonym of Gyrostomum Fr. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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