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Chacoan regions of Argentina (Provinces of Corrientes, Chaco, Formosa, Jujuy, Salta, Santiago del Estero, Santa Fe and Tucumán), Bolivia and Paraguay (Departments of Alto Paraguay, Boquerón, Central, Concepción, Presidente Hayes) , and open formations near of the rivers Beni, Madeira, Tapajos and Xingu at Amapá State, Brazil.


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