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BiologyFound in the sea, estuaries, and lower reaches of rivers (Ref. 4833). Commonly found on muddy shores in estuaries and in the tidal zone of rivers (Ref. 12693). Lives in burrows in the mud and emerges on sunny days at low tide. Moves quickly across the mud, but can be caught with entangling nets strung above the ground or with cast-nets tossed onto the exposed mud (Ref. 12693). An amphibious air-breather that actively shuttles back and forth between rock pools and air (Ref. 31184).