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BiologyAn intertidal species (Ref. 31184) found on sand and mud flats along bay shores. Also in estuarine areas, swamps, marshy areas and on tidal mud flats (Ref. 5258). Also found in the freshwater tidal zone (Ref. 12693). Actively shuttling back and forth between rock pools and air (Ref. 31184). Amphibious air-breather (Ref. 31184).