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BiologyOccurs in water bodies on low-lying plains, with little current (Ref. 26100). Found in a variety of habitats, most abundant in large rivers and streams with slow to moderate current. Avoids upper, cold water streams. Widely distributed and locally abundant, but populations are greatly fragmented and several of them are potentially threatened as many small streams turn dry during summer. Maximum life span is about 18 years (Ref. 59043).