Occurs in the middle reaches of streams with moderate current and abundant aquatic vegetation (Ref. 59043) and in the lower reaches of rivers with slow current and abundant submerged plants (Ref. 26100). Feeds on invertebrates, plants (Ref. 26100) and zooplankton (Ref. 59043). Spawns from March to May. Threatened due to water abstraction and the introduction of other species (Ref. 26100).