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BiologyInhabits silty backwaters of rivers and spawns in gravel riffles (Ref. 86798). Both the larval and adult stages were originally collected from the Friant-Kern Canal, which is 85% concrete lined and has a discharge greater than 57 m3/sec; an artificial habitat. Occurs in the Merced River, which should be considered typical habitat. Adults nonparasitic (Ref. 89241).