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 Life habit: lichenized; Thallus: foliose, c. circular in outline, lobate, contiguous, rarely imbricate; lobes: sublinear to subirregular; lobe tips: usually rotund, with bulbate cilia; upper surface: gray, smooth, plane to rugulose, shiny or dull, maculate or not, usually epruinose; with or without isidia, soredia or lobules; pseudocyphellae: absent; upper cortex: pored epicortex, palisade parenchymatous,; medulla: white, loosely packed; cell walls: containing isolichenan; photobionts: primary one a Trebouxia, secondary photobiont absent; lower surface: tan to black, plane to sometimes wrinked, attachment by simple and/or dichotomous rhizines; cyphellae, pseudocyphellae and tomentum absent; Ascomata: apothecial, laminal, orbicular, cup-shaped, subpedicellate or sessile; margin: prominent, with thalloid rim; disc: imperforate, pale to dark brown, coronate with black bulbae around inner margin of thalline exciple or ecoronate; exciple: gray or hyaline; epithecium: brown or brownish yellow; hypothecium: hyaline,; asci: lecanoral, wall layers apex thickened; apex: amyloid, with wide, axial body divergent towards apex, c. 8-spored; ascospores: simple, ellipsoid or bicornute, curved or semilunate, obtuse at the poles; 6-21 x 4-12 µm; walls: thin, hyaline, not amyloid; Conidiomata: absent or present, pycnidial, laminal or marginal, immersed or emergent sessile; conidia: bifusiform or rarely bacilliform, 5-10 (-15) x 1 µm; Secondary metabolites: upper cortex with atranorin (rarely usnic acid accessory) and chloroatranorin; medulla with some combination of orcinol depsides or depsidones, ß-orcinol depsides or depsidones; Geography: predominately pantropical and subtropical with a few species extending to temperate forests and woodlands; Substrate: mostly bark or acidic rock. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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