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BiologyFound in inshore habitats, lagoons, lakes and large to medium-size rivers; on sand or mud bottom. Spawns after first winter in March-July; individual females spawn several times during a season. Adhesive eggs deposited under or between stones, shells and aquatic plants and males guard the eggs until hatching. Feeds on a wide variety of benthic invertebrates (worms, amphipods, Ref.4696) (Ref.59043).