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BiologyInhabit middle and lower reaches, in deep and slow water, with abundant submerged plants (Ref. 11240). Feed on invertebrates, plants and fish (Ref. 26100). Juveniles prey on invertebrates while large adult individuals are mainly piscivorous. Spawn probably in May-June (Ref. 59043). Threatened by pollution, habitat destruction, introduction of new species (Ref. 26100) and water abstraction (Ref. 59043). Attains up to about 80 cm SL (Ref. 59043).