- Lima, F.C.T. 2003 Characidae - Bryconinae (Characins, tetras). p. 174-181. In R.E. Reis, S.O. Kullander and C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil. (Ref. 38504)
- Pearson, N.E. 1937 The fishes of the Atlantic and Pacific slopes near Cajamarca, Peru. Proceedings of the California Academy of Sciences, 4 (23) (7):87-98, pls. 12-13. (Ref. 38908)
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