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BiologyOccasionally form schools; mainly diurnal. Prefer shallow, vegetated areas (Ref. 4967). Fry are common in marginal vegetation and juveniles are found in the seasonal floodplain. Herbivorous, feed on water plants and epiphyton, and some invertebrates. Substrate spawner (Ref. 28714). Larvae develop in close association with substrate. Extended temperature range 6.5 - 42.5 °C, natural temperature range 10.5 - 36°C (Ref. 3).