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BiologyOccurs in both sediment-free and sediment-rich rocky biotopes. Favors clear, running waters in rocky habitats of small and large rivers, also found in lakes and dams over rocky areas. Feeds on diatoms and other small algae from the rocks (Ref. 5595). Also feeds on `aufwuchs' from the surface of rocks, tree trunks and other firm surfaces. Masses upstream in masses to breed, using the mouth & broad pectorals to climb damp surfaces of barrier rocks & weirs (Ref. 7248).