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Global Range: (20,000-2,500,000 square km (about 8000-1,000,000 square miles)) Range includes North American Gulf Coast drainages from Halawakee Creek in the Chattahoochee River system in east-central Alabama to Neches River system in Texas; and north in the Mississippi River drainage to northern Mississippi and Ouachita river system, Arkansas; disjunct populations in Bear Creek system, tributary to the Tennessee River in northwestern Alabama and northeastern Mississippi (Retzer et al. 1986, Mettee et al. 1996, Piller et al. 2001, Page and Burr 2011).