Biology

Most of the specimens were caught in a flooded/marshy area draining into the stream, with little current on a muddy bottom with a large amount of decomposing litter. Few specimens were caught in the stream itself, in a margin area of slower current under riparian vegetation over a bottom of rocks, pebbles and sand, with submerged litter only on the bottom of the larger and deeper pools. The species eats mainly aquatic and terrestrial arthropods. Sexual dimorphism (Ref. 52251).