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BiologyFound in lowland water bodies, with little current, and in springs and associated wetlands (Ref. 26100). Inhabits streams and sources with clear (often cold) water. Occurs in subterranean waters during winter or droughts. Threatened due to pollution, habitat destruction and introduction of Alburnus arborella, Squalius cephalus and Lepomis gibbosus (Ref. 59043).