Bathygobius ramosus — Details

Panamic Frillfin learn more about names for this taxon

Overview

Comprehensive Description

Description

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Common names: frillfin (English), gobio (Espanol), mapo (Espanol)
 
Bathygobius ramosus Ginsburg, 1947


Panamic frillfin


Body robust; head broad, depressed; snout bluntly rounded; eyes large, close together; mouth moderate sized, relatively horizontal; tongue with distinct notch in front; jaw teeth in bands, without canines; dorsal fins VI + I, 8-10 rays; anal I, 7-9 rays; pelvic disc rounded, front membrane with large side lobes; tail round, 24-31 rays; pectoral rays 18-21, the uppermost 6 rays forming free filaments, usually the 1st  and invariably the remaining five branching more than once; scales large, rough, 32-38 along midside, predorsal scales not reaching eyes, remainder of head scaleless.

Olive green to brown with lighter scale centres; often a row of dark spots along midside and diffuse bars or saddle may be present on back; lower head with small white ocelli; first dorsal fin entirely pigmented, and with a central stripe; caudal and second dorsal fin with wavy brown bands or spots; 1st  of 3 dark blotches behind eye is largest.


Size: grows to 11.4 cm.

Inhabits the rocky intertidal zone.

Depth: 0-2 m.

Southern Baja and the Gulf of California to northern Peru, Cocos Island, and the Revillagigedos Islands. According to Miller et al 2001 there are two subspecies - Bathygobius r. longipinnis from the Revillagigedos and Tres Marias, and Bathygobius r. ramosus from the remainder of the range.

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Biology

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Inhabits high, sandy-bottom tide pools along rocky coasts.
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Distribution

Range Description

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This species is endemic to the Eastern Pacific, ranging from southern Baja and the Gulf of California to northern Peru, including Cocos Island, and the Revillagigedo and Tres Marias Islands of Mexico.
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Depth

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Depth Range (m): 0 (G) - 2 (G)
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Zoogeography

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See Map (including site records) of Distribution in the Tropical Eastern Pacific


 
Global Endemism: All species, East Pacific endemic, Tropical Eastern Pacific (TEP) endemic

Regional Endemism: All species, TEP endemic, Continent + Island (s), Continent, Island (s)

Residency: Resident

Climate Zone: North Temperate (Californian Province &/or Northern Gulf of California), Northern Subtropical (Cortez Province + Sinaloan Gap), Northern Tropical (Mexican Province to Nicaragua + Revillagigedos), Equatorial (Costa Rica to Ecuador + Galapagos, Clipperton, Cocos, Malpelo), South Temperate (Peruvian Province )

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Eastern Pacific: Bahía Magdalena and the Gulf of California in Mexico to northern Peru.
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Physical Description

Morphology

Size

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Length max (cm): 11.4 (S)
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Size

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Maximum size: 114 mm TL
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Max. size

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20.0 cm TL (male/unsexed; (Ref. 55763))
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Diagnostic Description

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Body small; tip of tongue bifurcated; two dorsal fins, the first with 7 flexible spines; scales large and readily visible; tips of uppermost 3 to 5 pectoral rays free; body olive green to brown; flanks with dark spots and/or diffuse (Ref. 55763).
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Type Information

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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Dow
Locality: Central America, W. Coast C.A., Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Holotype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Balboa, Canal Zone., Panama, Pacific
  • Holotype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): G. Miller
Year Collected: 1937
Locality: Venado Beach, Panama, Canal Zone, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): Ray & Don
Year Collected: 1935
Locality: Pinas Bay, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): W. Schmitt
Year Collected: 1935
Locality: Secas Isle. Outer Side Small Most Western Island, Tidepools, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Zetek
Year Collected: 1914
Locality: Taboga Island, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Zetek
Year Collected: 1914
Locality: Taboga Id, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Zetek
Year Collected: 1914
Locality: Taboga Id, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Zetek
Year Collected: 1914
Locality: Panama City, Panama Province, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Year Collected: 1904
Locality: Perico Island, Panama, C. Z., Pool, Panama, Pacific
Vessel: Albatross
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools Panama, Panama., Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Balboa, Canal Zone, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): C. Gilbert & Leland Stanford Junior University
Locality: Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Panama City, Panama., Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tidepools, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Panama, Panama., Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Panama City, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1911
Locality: Taboga Island, Panama, Taboga Island, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Naos Island, C.Z., Canal Zone, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Balboa, C. Z., Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Meek & S. Hildebrand
Year Collected: 1912
Locality: Tide Pools, Balboa, Canal Zone., Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): A. Busck
Year Collected: 1907
Locality: Tobago Island, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Morrison & A. Wetmore
Year Collected: 1944
Locality: Pearl Islands: San Jose Is., Intertidal Zone of S. Side of Small E. Bay, Panama, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): J. Morrison & Wetmore
Year Collected: 1944
Locality: R. De P.: Pearl Islands, San Jose Island, Tidepools On N. Side of Small E. Bay, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): A. Wetmore & J. Morrison
Year Collected: 1944
Locality: Panama, Pearl Islands: San Jose Is., Mouth of Stream S. of Rio Marina (Brackish Water), Panama, North America
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Panama: San Francisco Reef (Tide Pools), Panama City, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Taboga Island, Panama Bay, Tidepool, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Pearl Islands, R.P. (Saboga Island), (Tide Pools), Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Panama: Venado Beach (Pacific), Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Near Panama City, San Francisco Beach, tide pools, Panama Province, Panama, Gulf of Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Pacific Coast of Panama, West Coast, Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Pearl Islands, San Jose Island, North Side of Bay, San Jose Island, Panama, Pearl Islands, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Panama: Pearl Islands, Panama, Pearl Islands, Gulf of Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): S. Hildebrand
Year Collected: 1937
Locality: Panama: Pearl Islands, San Jose Island, San Jose Island, Panama, Pearl Islands, Gulf of Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Paratype for Bathygobius ramosus ramosus Ginsburg
Collection: Smithsonian Institution, National Museum of Natural History, Department of Vertebrate Zoology, Division of Fishes
Collector(s): Foster
Year Collected: 1937
Locality: Bella Vista, R. P., Panama, Pacific
  • Paratype: Ginsburg, I. 1947. Journal of the Washington Academy of Sciences. 37 (8): 282.
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Ecology

Habitat

Habitat and Ecology

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Habitat and Ecology
This shallow-water species is found in sandy-bottom tide pools to depths of 2m along rocky coasts. In Gorgona Island, Colombia (Rubio, 1986), this species is frequently found in coraline substrate. It is a carnivore, feeding on mobile benthic worms, crustaceans and gastropods.

Systems
  • Marine
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Depth range based on 64 specimens in 1 taxon.
Water temperature and chemistry ranges based on 11 samples.

Environmental ranges
  Depth range (m): 0.16 - 25.62
  Temperature range (°C): 21.311 - 27.666
  Nitrate (umol/L): 0.144 - 1.599
  Salinity (PPS): 32.938 - 34.642
  Oxygen (ml/l): 4.671 - 5.074
  Phosphate (umol/l): 0.327 - 0.429
  Silicate (umol/l): 2.028 - 3.286

Graphical representation

Depth range (m): 0.16 - 25.62

Temperature range (°C): 21.311 - 27.666

Nitrate (umol/L): 0.144 - 1.599

Salinity (PPS): 32.938 - 34.642

Oxygen (ml/l): 4.671 - 5.074

Phosphate (umol/l): 0.327 - 0.429

Silicate (umol/l): 2.028 - 3.286
 
Note: this information has not been validated. Check this *note*. Your feedback is most welcome.
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Habitat

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Salinity: Marine, Brackish

Inshore/Offshore: Inshore, Inshore Only

Water Column Position: Bottom, Bottom only

Habitat: Reef (rock &/or coral), Rocks, Reef and soft bottom, Reef associated (reef + edges-water column & soft bottom), Soft bottom (mud, sand,gravel, beach, estuary & mangrove), Estuary, Mangrove

FishBase Habitat: Reef Associated
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Environment

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reef-associated; marine
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Trophic Strategy

Feeding

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Feeding Group: Carnivore

Diet: mobile benthic worms, mobile benthic crustacea (shrimps/crabs), mobile benthic gastropods/bivalves
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Life History and Behavior

Reproduction

Reproduction

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Egg Type: Benthic, Pelagic larva
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Molecular Biology and Genetics

Molecular Biology

Statistics of barcoding coverage: Bathygobius ramosus

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Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Species: 9
Species With Barcodes: 1

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Conservation

Conservation Status

IUCN Red List Assessment

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Red List Category
LC
Least Concern

Red List Criteria

Version
3.1

Year Assessed
2010

Assessor/s
Findley, L., Van Tassell, J.

Reviewer/s
Carpenter, K., Polidoro, B., Livingstone, S. (Global Marine Species Assessment Team)

Justification
This species is widespread and abundant in the Eastern Pacific. There are no major threats known to this species, and no current indication of widespread population decline. It is listed as Least Concern.
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Conservation status

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IUCN Red List: Not evaluated / Listed

CITES: Not listed
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Population

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Population
This species is very common and abundant throughout its range within suitable rocky intertidal habitat.

Population Trend
Stable
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Threats

Threats

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Major Threats
There are no major threats known for this species.
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Least Concern (LC)
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Management

Conservation Actions

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Conservation Actions
There are no known conservation measures for this species. However, this species distribution falls partially into a number of Marine Protected Areas in the Eastern Pacific region (WDPA 2006).
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Relevance to Humans and Ecosystems

Benefits

Importance

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aquarium: commercial
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