Biology

A resident intertidal species with homing behavior (Ref. 32612, 48637) and amphibious air-breather (Ref. 31184, 79840) found in brackish mud flats in mangrove and nipa palm areas. Occasionally in the lower parts of freshwater streams (Ref. 2847, 44894, 48637, 79840). Actively shuttling back and forth between rock pools and air (Ref. 31184). Feed on worms, crustaceans, and insects (Ref. 37816). Can stay out of the water for up to 37 h if kept moist (Ref. 51276).