Biology

Occurs in rocky rapids and cascades in altitudes of 900-1,200 m. Prefers deeper waters (30-60 cm) over cobbles (Ref. 26172). Feeds on benthic invertebrates such as mayfly and caddis fly nymphs, blackfly and midge larvae and small snails. Oviparous (Ref. 205). Breeds during summer and is a partial spawner. Increasing extraction and regulation of river water as well as pollution threaten this species.