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BiologyFound in relatively deep (1-2 m) fast-flowing water, in areas with gravel or sand substrates. Juveniles (1-3cm TL) shoal together in schools of 30-100 individuals in very shallow water (5-25 cm), downstream. Adults occupy deeper water (1-2m), upstream. Usually occupies unshaded areas. Fast swimmers and not easily identified from water surface. Declining numbers due to intensive fishing by fish exporters.