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BiologyInhabits piedmont and mountain streams with Mediterranean water regime, strong to moderate current, rock or gravel bottom. Abundant along shores and may be restricted to very small pools during summer. Feeds predominantly on terrestrial insects, benthic invertebrates and algae. Breeds in shallow riffle habitats, with strong current and stone or pebble bottom, at 10-20 cm water depth. Moves downstream in winter to deeper pools and to uppermost stretches of streams in summer (Ref. 59043).