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BiologyFound in rivers, reservoirs, lakes and canals (Ref. 11145). Occurs in the sea at depths of up to 25 m and in rivers in deep stretches with gravel or stone bottom. In freshened parts of sea, preys mainly on benthic crustaceans. Does not feed while migrating upstream and starts to feed again at the spawning sites mainly on insects, juveniles of other fishes, rarely on algae, seeds and other plant material. Spawns in fast-flowing water at areas with hard bottom and depths of 1-2 m (Ref. 59043).