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BiologyAdults inhabit upper edge of outer reef slopes and inshore rocky reefs. Juveniles associated with drifting seaweed (Ref. 12114, 12115). Benthopelagic (Ref. 58302). Feed on zooplankton, benthic algae, and small invertebrates (Ref. 1602). Often in aggregations (Ref. 9710) feeding at midwater or tending nests among rocks and coral ledges (Ref. 90102). In large numbers at spawning sites that are timed with large tides that carry their pelagic offspring far offshore (Ref. 48636). Oviparous, distinct pairing during breeding (Ref. 205). Eggs are demersal and adhere to the substrate (Ref. 205). Males guard and aerate the eggs (Ref. 205).