Comprehensive Description

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 Perennial woody herbs or subshrubs20–50 cm tall, stems prostrate or sprawling to erect, branching from near base, with sparse lateral branches, glabrous or new growth sparsely pilose, mature twigs 2–3 mm diam, bark smooth, reddish brown to dark brown, with tufts of pilose hairs at thickened, persistent leaf bases.Leavessimple, spirally arranged; blade dark green above, yellow-green below, firm and moderately coriaceous when fresh, chartaceous when dry, glabrous, (15–) 20–47 × (12–)18–37 mmovate to broadly ovate, base obtuse to rounded or truncate, apex obtuse, tip usually emarginate, venation subpalmate with 1–2(–3) pairs of secondary veins from base and 2–3 pairs along midrib above, venation raised and visible to 3° above and to 4° beneath; margin thin, plane; petiole (20–) 25–45 × 0.6–0.8 mm, sparsely scattered pilose or glabrescent, flattened, adaxially sulcate, distally with slight pulvinus.Inflorescencesaxillary near ends of branches, cymose, 5–13–flowered, 5–8 cm long, peduncles 4–5 cm long, terminating in apical flower and two monochasial lateral branches 18–40 mm long each with 2–6 flowers, bracts linear-subulate, 1–2 × 0.3–0.4 mm, sparsely puberulent with acicular and scattered glandular-tipped trichomes.Flowers(long-styled morph seen) with 5 calyx lobes 5–7 × 1.7–2 mm, narrowly ovate-oblong, 6–8-veined, apex acute, both surfaces minutely glandular-puberulent with capitate trichomes; petals yellow, 10–12 × 5 mm, narrowly obovate to oblong-elliptic, 7–9-veined, apex obtuse to rounded; stamens 10, in two series, the longer 7–8 mm long, the shorter 5.5–6 mm long, filaments connate basally, anthers broadly ellipsoid, 0.4–0.5 mm long, reniform to subcircular; gynoecium 10-14 mm long, with ovary 6–7 mm long, narrowly ovoid-cylindrical, externally glabrous, beak 4–5 mm long, styles 5, 1–2.5 mm long, stigmas slightly thickened, papillose, not bifid.Fruitsovoid-cylindrical, 10–11 × 2–2.5 mm, at maturity apparently straight and not twisting, externally glabrous, carpels 5, villous within, seeds ca. 40.Seedsellipsoid, compressed, 0.8–0.9 × 0.5 mm, surface shiny, brown, rugose.


Creative Commons Attribution 3.0 (CC BY 3.0)

© David H. Lorence, Warren L. Wagner

Source: PhytoKeys

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