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BiologyInhabit rocky pools of headwaters, creeks and small rivers, and rocky margins of medium-sized rivers and impoundments (Ref. 5723); also found in streams (Ref. 10294). Adults feed on midge, mayfly, and immature caddisfly, with copepods and other microcrustaceans (Ref. 10294). Eggs are found clustered on underside of stone and guarded by males (Ref. 7043).