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BiologyAmphibious air-breather (Ref. 31184) that skips or walks on sand or mud in search of food; that is, on exposed intertidal mud flats of estuarine mangrove swamps. Adult feed chiefly on arthropods (crabs, insects, etc.) of the mud surface. Also included in the diet is the white mangrove, Avicennia nitida (Ref. 3026). Spawn in burrows (Ref. 3657).