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BiologyAdults occur in rubble and gravel riffles, less often in sand-gravel runs of headwaters, creeks and small rivers. Also found in springs and their effluents and rocky shores of lakes (Ref. 1998, 10294). Feed mainly on aquatic insect larvae, but also on crustaceans, annelids, fishes, fish eggs, and plant material (Ref. 1998, 10294). Spawn in the spring (Ref. 1998). A male guards the cluster of eggs laid by different females (Ref. 1998). Neither anterolateral glandular groove nor venom gland is present (Ref. 57406).