The Nile perch (Lates niloticus) is a species of freshwater fish native to the Congo, Nile, Senegal, Niger, and Lake Chad, Volta, Lake Turkana and other river basins, and is now widespread through tropical Africa. It has a large number of common names including African snook, Capitaine, Victoria perch and many local names in various African languages, such as the Luo name Mbuta. One of the largest freshwater fish, Lates niloticus reaches a maximum length of nearly two meters (more than six feet), weighing up to 200 kg. Mature fish average 121–137 cm although many fish are caught before they can grow this large.
A fierce predator that dominates its surroundings, the Nile perch feeds on fish (including its own species), crustaceans, and insects; the juveniles also feed on zooplankton. Nile perch have been introduced to many other lakes in Africa, including Lake Victoria in 1962. The Lake Victoria introduction is an often-cited example of enormous effect of a non-native species upon its new surroundings, as Nile perch decimated the rich diversity of hundreds of native species, and caused the decline or extinction of an estimated 200 chichlid fish in Lake Victoria. This highly studied introduction caused a booming fishing industry for Nile Perch which destroyed the livelihood of traditional local lake-side dwelling people and caused a chain of other high impact repercussions on the environment and economy of the area. The IUCN's (World Conservation Union) Invasive Species Specialist Group considers Lates niloticus one of the world's 100 worst invasive species. The state of Queensland in Australia levies heavy fines on anyone found in possession of a living Nile perch, since it competes directly with the native Barramundi, which is similar but does not reach the same size as the Nile perch.
The species is of great commercial importance as a food fish. The Nile perch is also popular with sport anglers as it attacks artificial fishing lures and is also raised in aquaculture.
(CABI 2011; Lipton 2003; Schofield 2012; Wikipedia 2012)
Lates niloticus (Nile perch) is widespread throughout the Ethiopian Region of Africa, occurring commonly in all major river basins including Nile, Chad, Niger, Senegal and Volta. The nilotic population penetrates northwards well into the geographical limits of the Mediterranean Region and is present in the waters of Lake Mariout situated in the Nile Delta. Southwards the distribution includes parts of the Congo Basin. The most common place to find the Nile perch is in Lake Victoria where the species was introduced in 1962.
Biogeographic Regions: nearctic (Introduced ); ethiopian (Introduced , Native )
- Hopson, A. 1972. A Study of the Nile Perch (*Lates niloticus* (L.) Pisces: Centropomidae) in Lake Chad. Foreign and commonwealth office overseas development administration (Overseas Research Publication), 19: 1-90.
Central Africa: In Lower Guinea it is known from the Sanaga and Cross and coastal rivers of Cameroon. It has been introduced, at unknown date, from Sudan to Congo. According to FAO (2005), it is naturally reproducing and established. However, the species is native in the Congo River basin. Except for records from its natural distribution within the Lower Guinea province, no museum records are available.
Eastern Africa: This species is present in Lake Albert, the Albert and Murchison Niles and Turkana. It is now fully established in Lakes Victoria, Kyoga and Nabugabo, and the Victoria Nile, through introductions. According to Hartley (1984) an unpublicised introduction of L. niloticus took place in Lake Naivasha in the early 1970s and since the early 1980s several perch have been caught. No information is available on its present status but probably the species did not establish in the lake.
Northern Africa: It is common in the Delta, Lower and This species is known from upper Egyptian Nile, as well as Lakes Wadi El-Rayan and Burollos, and Nozha Hydrodrome. It is present in the brackish waters of Lake Mariout, near Alexandria.
Northeast Africa: It is found throughout the Nile drainage, Lakes Chamo and Abaya as well as Baro and Tekeze basins in Ethiopia. Also Setit in Eritrea.
Western Africa: This species is found almost everywhere in West Africa. Widely distributed in the Volta basin. (Dankwa et al. 1999) Present in Black Volta, White Volta and the Oti (Dankwa 1984).
Nile perch are silver in color with a blue tinge. They have a distinctive dark black eye with a bright yellow outer ring. Nile perch are usually seen around 2-4 kg, but have been caught and seen at sizes up to 200 kg (the largest at 232 kg). They average around 85-100 cm but can grow to 193 cm. The females are generally larger than males. The preopercle and pre-orbital bones are armed with spines, with a large spine on the free edge of the operculum.
Range mass: 232 (high) kg.
Average mass: 2-4 kg.
Range length: 193 (high) cm.
Average length: 85-100 cm.
Other Physical Features: ectothermic ; heterothermic ; bilateral symmetry
Sexual Dimorphism: female larger
Nile perch are found in many different types of fresh water. They prefer warm, tropical waters (27°N – 7°S) where they grow to large sizes and occur in high densities. Adult Nile perch occupy all habitats in lakes and rivers (10-60 m in depth) where there is enough oxygen with the exception of rocks, swamps, and the pelagic zone. Small juveniles are restricted to shallow near-shore environments (Luna, 2002; Queensland Government, 2002).
Range depth: 0 to 60 m.
Habitat Regions: tropical ; freshwater
Aquatic Biomes: lakes and ponds; rivers and streams
- FishBase, 2004. "Lates niloticus, Nile perch" (On-line). FishBase. Accessed 06/08/04 at http://www.fishbase.org/Summary/SpeciesSummary.cfm?genusname=Lates&speciesname=niloticus.
- Queensland Department of Primary Industries and Fisheries, 2004. "Fish - Note, Nile Perch (Lates niloticus)" (On-line). Accessed November 1, 2002 at http://www.dpi.qld.gov.au/fishweb/2374.html.
Habitat and Ecology
Introductions in Lake Victoria were mainly from Lake Albert. Nile perch is responsible through predation and competition for food of the decimation and possible disappearance of two hundred or more species of the unique flock of endemic haplochromine cichlids in Lake Victoria.
From 10 to 29 meters.
The diet of Nile perch consists of fishes, insects, crustacea and mollusks. The type of prey ingested by the predator depends on the predator size, prey availability and abundance within a given habitat (Ogari, 1984). Juvenile Nile perch feed on copepods, prawns in the genus Caridina, fish fry, small gastropods, and bivalves. As the fish matures and moves to greater depths haplochromine cichlids constitute over 95% of their food consumption. Occasional items found in the Nile perch's diet include smaller fish in the genera Barbus, Clarias, Haplochromis, Lates, Oreichromis, and Xenoclarias. Besides crustacean zooplankton, invertebrate prey includes snails, clams, and insects (odonate larvae, aquatic Hemiptera, mayflies in the genus Povilla, and larvae of phantom midges (Chaoborus). Fish in the genus Rastrineobola are very common in the diet in terms of occurrence, and are second to haplochromines (Acere, 1985). As Nile perch grow larger, they take larger prey. Nile perch less than 80 cm tend to feed on smaller fishes than those greater than 80 cm. This demonstrates that the predator is capable of shifting to other sizes of prey when more suitable sizes become scarce (Ogutu-Ohwayo, 1984).
Animal Foods: fish; insects; mollusks; aquatic crustaceans; zooplankton
Primary Diet: carnivore (Piscivore )
- Ogari, J. 1984. Distribution, Food and Feeding Habits of *Lates niloticus* in Nyanza Gulf of Lake Victoria (Kenya). FAO Fisheries Report, 335: 68-80.
- Ogutu-Ohwayo, R. 1984. The Effects of Predation by Nile Perch, *Lates niloticus* (Linne) Introduced into Lake Kyoga (Uganda) in Relation to the Fisheries of Lake Kyoga and Lake Victoria. FAO Fisheries Report, 335: 18-41.
Fish Species Associates in the Senegal River
There are 141 species of fish recorded in the Senegal River, most of which are native; however, there are no endemic species of fish in the Senegal Basin. Among the larger native benthopelagic taxa are: the 170 centimetre (cm) North African Catfish (Clarias gariepinus), the 149 cm Electric Catfish (Malapterurus electricus), and the 92 cm African Carp (Labeo coubie).
Some of the larger native demersal fishes of the Senegal Basin are: the 204 cm Aba (Gymnarchus niloticus). the 200 cm Nile Perch (Lates niloticus), the 183 cm Sampa (Heterobranchus longifilis), and the 150 cm Cornish Jack (Mormyrops anguilloides).
Pelagic native fishes in the Senegal River include the 65 cm True Big-scale Tetra (Brycinus macrolepidotus) and the 16 cm Ansorge Fangtooth Pellonuline (Odaxothrissa ansorgii)
The Nile perch acts as a major predator in its native and introduced habitats.
Nile perch have been observed with several different kinds of parasites, Lernea (region after the operculum), arguilids (in the gills), and various nematodes (throughout the body).
Known prey organisms
This list may not be complete but is based on published studies.
Diseases and Parasites
Life History and Behavior
Perception Channels: tactile ; chemical
The growth of the Nile perch is very fast during the first year. The rate then decreases during the second, third, fourth and fifth years.
- Acere, T. 1984. Observations on the biology, age, growth, maturity and sexuality of Nile perch, *Lates niloticus* (Linne), and the growth of its fishery in the northern waters of Lake Victoria. FAO Fisheries Report, 335: 42-61.
Nile perch live up to the age of 16 years. There is a higher mortality rate for males than females.
Status: wild: 16 (high) years.
Status: captivity: 7.8 years.
Nile perch are sexually dimorphic. The male has only anal and urogential openings just anterior to the anal fin, whereas the female has a genital orifice separate from the urinary opening. They become sexually mature at the age of 3 years. Males dominate the sex ratio up to 80 cm TL, while the females are dominate at 80 cm TL and above (Asila and Ogari, 1988). Ovulation takes place in the spring with the rising water temperature. Spawning is usually done in sheltered areas, but can also occur in open waters.
Breeding season: Breeding peaks from March to June.
Range number of offspring: 3,000,000 to 15,000,000.
Average number of offspring: 9,000,000.
Average gestation period: 20 hours.
Average age at sexual or reproductive maturity (female): 3 years.
Average age at sexual or reproductive maturity (male): 3 years.
Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; fertilization (External ); oviparous
Average age at sexual or reproductive maturity (female)
Sex: female: 912 days.
Parental Investment: no parental involvement
- Hopson, A. 1972. A Study of the Nile Perch (*Lates niloticus* (L.) Pisces: Centropomidae) in Lake Chad. Foreign and commonwealth office overseas development administration (Overseas Research Publication), 19: 1-90.
Molecular Biology and Genetics
Barcode data: Lates niloticus
Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species.
See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
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Download FASTA File
Statistics of barcoding coverage: Lates niloticus
Public Records: 7
Specimens with Barcodes: 131
Species With Barcodes: 1
IUCN Red List of Threatened Species: least concern
IUCN Red List Assessment
Red List Category
Red List Criteria
Relevance to Humans and Ecosystems
The introduction of the Nile Perch to Lake Victoria has caused serious ecological problems. The richness and diversity of endemic cichlid species is rapidly declining. Over 300 native species have already been driven to extinction due to the feeding patterns of the Nile perch (Schofield, 1999). Although for the time being the strong increase of L. niloticus seems to be a favorable development for the fishing industry, the final consequences may be very serious for future fish production in the lake (Goudswaard and Witte, 1984). Since the increase of Nile perch, the accelerated decline in diversity has altered the food web structure and caused ecological changes due largely to human actions, which have profound socioeconomic effects (Kitchell and Schindler, 1997). The continuing degradation of Lake Victoria's ecological functions has serious long-term consequences for the ecosystem services it provides and may threaten social welfare in the different countries bordering its shores (Verschuren and Johnson, 2002). Also since the increase of Nile perch, smaller scale fishing companies have been hurt significantly.
- Goudswaard, P., F. Witte. 1984. Observation on Nile perch, *Lates niloticus* (L.), 1758, in the Tanzanian waters of Lake Victoria. FAO Fisheries Report, 335: 62-67.
- Schofield, P. 1999. Interactions between Nile perch, *Lates niloticus*, and other fishes in Lake Nabugabo, Uganda. Environmental Biology of Fishes, 55 (4): 343-358.
- Verschuren, D., T. Johnson. 2002. History and timing of human impact on Lake Victoria, East Africa. Proceedings of the Royal Society Biological Sciences Series B, 269 (1488): 289-294.
The Nile perch has yielded an increase in total fishery, and fishery-related employment has increased dramatically. Since the increase of Nile perch, larger factory fishing companies have thrived greatly.
Positive Impacts: food
- Kitchel, J., D. Schindler. 1997. The Nile perch in Lake Victoria: Interactions between predation and fisheries. Ecological Applications, 7 (2): 653-664.
The Nile perch (Lates niloticus) is a species of freshwater fish in family Latidae of order Perciformes. It is widespread throughout much of the Afrotropic ecozone, being native to the Congo, Nile, Senegal, Niger, and Lake Chad, Volta, Lake Turkana, and other river basins. It also occurs in the brackish waters of Lake Maryut in Egypt. Originally described as Labrus niloticus, among the marine wrasses, the species has also been referred to as Centropomus niloticus. Common names include African snook, Victoria perch (a misleading trade name, as the species is not native to Lake Victoria), and a large number of local names in various African languages, such as the Luo name mbuta or mputa. In Tanzania, it is called sangara, sankara or chenku. In Francophone African countries, it is known as capitaine and in Egypt/Sudan as am'kal. Its name in the Hausa language is giwan ruwa, meaning "water elephant".
Lates niloticus is silver in colour with a blue tinge. It has distinctive dark-black eyes, with a bright-yellow outer ring. One of the largest freshwater fish, it reaches a maximum length of nearly 2 m (more than 6 ft), weighing up to 200 kg (440 lb). Mature fish average 121–137 cm (48–54 in), although many fish are caught before they can grow this large.
Adult Nile perch occupy all habitats of a lake with sufficient oxygen concentrations, while juveniles are restricted to shallow or nearshore environments. A fierce predator that dominates its surroundings, the Nile perch feeds on fish (including its own species), crustaceans, and insects; the juveniles also feed on zooplankton. Nile perch use schooling as a mechanism to protect themselves from other predators.
Nile perch have been introduced to many other lakes in Africa, including Lake Victoria (see below) and the artificial Lake Nasser. The IUCN's (World Conservation Union) Invasive Species Specialist Group considers L. niloticus one of the world's 100 worst invasive species.
The state of Queensland in Australia levies heavy fines on anyone found in possession of a living Nile perch, since it competes directly with the native barramundi, which is similar and grows to 1.8m long while the Nile Perch grows to 2m long.
Lake Victoria introduction
The introduction of this species to Lake Victoria is one of the most cited examples of the negative effects alien species can have on ecosystems.
The Nile perch was introduced to Lake Victoria in East Africa in the 1950s, and has since been fished commercially. It is attributed with causing the extinction or near-extinction of several hundred native species, with some populations fluctuating with commercial fishing and the actual Nile perch stocks. The Nile perch initially fed on native cichlids, but with decreasing availability of this prey, it now consumes mainly small shrimp and minnows.
The fish's introduction to Lake Victoria was ecologically disruptive, and lead to the establishment of large fishing companies. In 2003, Nile perch sales to the EU reached 169 million euros. Sport-fishing in the region of Uganda and Tanzania provided additional income from tourism. The long-term outlook is unclear, as overfishing is reducing L. niloticus populations.
The alteration of the native ecosystem had disruptive socioeconomic effects on local communities bordering the lake. Large-scale fishing operations, while earning millions of dollars from their exported L. niloticus catches, have displaced many local people from their traditional occupations in the fishing trade and brought them into the cash economy or - before the establishment of export-oriented fisheries - turned them into economic refugees. At least initially[verification needed], nets strong enough to hold adult Nile perch could not be manufactured locally and had to be imported for a high price.
The introduction of Nile perch has also had additional ecological effects on shore. Native cichlids were traditionally sun-dried, but because Nile perch have a high fat content (higher than cichlids), they need to be smoked to avoid spoiling. This has led to an increased demand for firewood in a region already hard-hit by deforestation, soil erosion, and desertification.
The Academy Award-nominated documentary Darwin's Nightmare by Hubert Sauper (a French-Austrian-Belgian production, 2004) deals with the damage that has been caused by Nile Perch introduction, including the import of weapons and ammunition in cargo planes from Europe, which are then used to export Nile perch, further exacerbating conflict and misery in the surrounding regions.
Regardless of whether it is considered positive or negative, the trophic web of Lake Victoria appears to have been drastically impoverished by the introduction of this novel near-top-level predator. While the lake ecosystem seems to be moving towards a new equilibrium, neither its former state nor the state of fisheries on Lake Victoria can ever easily be brought back.
On Lake Victoria, the only (small) trawlers present belong to research institutes. Small-scale fishing boats are propelled mostly by sails, and paddles are used on the smallest boats. However the number of boats propelled by outboard engines is on the rise, denoting a greater capital intensity of the local Nile perch fishery (see Beuving, 2013). One to three fishermen use a boat. The fish is caught mainly with gill nets and handlines and sometimes (short) longlines. Those caught by gill nets are usually dead when the nets are lifted. The fish are kept in the boat without protection or ice and taken to landing sites, mostly beaches, where they are weighed and purchased by company buyers using insulated boats or vans with ice, or the fish is bought by local women.
The fishery also generates indirect employment for additional multitudes of fish processors, transporters, factory employees, and others. All along the lakeshore, 'boom towns' have developed in response to the demands of fishing crews with money to spend from a day's fishing.[Note 1] These towns resemble shanties, and have little in the way of services. Of the 1,433 landing sites identified in the 2004 frame survey, just 20% had communal lavatory facilities, 4% were served by electricity, and 6% were served by a potable water supply.
Nile perch bought at the beach by women is usually cut into large pieces and smoke-dried for sale in distant places. Those bought by company buyers - usually company drivers - are placed on ice in an insulated company van or collection boat. After one to three days the van or boat will take the fish to a processing plant where the fish is filleted and the fillets are exported either by air if fresh or by boat if frozen. Local people around Lake Victoria prefer to eat tilapia rather than Nile perch, but in West Africa, Sudan, and Egypt, as well as in Israel, it is highly appreciated. In the 1990s, the value of Nile perch exports from Lake Victoria reached almost US$ 300 million per year.
The yield of fillets from a whole ungutted fish is about 30%. The remainder is head, skin, guts, bones and fins plus meat attached to the filleting frame. The frames used to be smoke-dried for local consumption, while heads and skins were used as fuel under frying pans to collect oil from the guts. Now, the companies process the filleting waste to fish meal. However, the swim bladder is dried and sold to traders for export to Southeast Asia where they are used as food.
- See for an anthropological study of these towns, called village landings, Beuving (2010).
- Azeroual, A., Entsua-Mensah, M., Getahun, A., Lalèyè, P., Moelants, T. & Ntakimazi, G. 2010. Lates niloticus. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.2. <www.iucnredlist.org>. Downloaded on 01 January 2014.
- Kaufman, Les. "Catastrophic Change in Species-Rich Freshwater Ecosystems: The lessons of Lake Victoria". BioScience 42 (11). doi:10.2307/1312084.
- Wood (1983). The Guinness Book of Animal Facts and Feats. Sterling Pub Co Inc. ISBN 978-0-85112-235-9.
- "Nile perch". Encyclopædia Britannica. Retrieved 2011-06-27.
- Pringle, Robert M. (2005-01-01). "The Nile Perch in Lake Victoria: Local Responses and Adaptations". Africa: Journal of the International African Institute 75 (4): 510–538. doi:10.3366/afr.2005.75.4.510. ISSN 0001-9720. JSTOR 3556959.
- Ben-Yami, M. 1996. Ecological and socioeconomic aspects of the expansion of Nile Perch in Lake Victoria. Pp. 95-110 in: Meyer, R.M. et al (Eds). Fisheries Resource Utilization and Policy. Proc. World Fisheries Congress. Theme 2. Oxford & IBH Publ.Co., New Delhi. 1996.
- J. Joost Beuving (2010). "Playing pool along the shores of Lake Victoria: Fishermen, careers and capital accumulation in the Ugandan Nile perch business". Africa 80 (2): 224–248. doi:10.3366/afr.2010.0203.
- LVFO (Lake Victoria Fisheries Organisation) (2005). Regional report on Lake Victoria Frame surveys for 2000, 2002 and 2004. Jinja, Uganda: LVFO and the Lake Victoria Environmental Management Project.
- Werimo, K.E.O. (1998). Nile perch oil characteristics, composition and use. FAO Expert Consultation on Fish Technology in Africa. 6, Kisumu (Kenya), 27-30 Aug 1996.
- Beuving, J. J. 2010. "Playing pool along the shores of Lake Victoria. Fishermen, careers and capital accumulation in the Ugandan Nile perch business" Journal of the International African Institute 80 (2): 224-248.
- Beuving, J. J. 2013. "Chequered Fortunes in Global Exports: The Sociogenesis of African Entrepreneurship in the Nile Perch Business at Lake Victoria, Uganda" ."
- Pringle, R.M. 2005. "The origins of the Nile perch in Lake Victoria." BioScience 55:780-787.
- Masciarelli, Alex. "The rise and fall of the Nile Perch." March 15, 2007. 
- Socio-economic effects of the evolution of Nile perch fisheries in Lake Victoria: a review. J. Eric Reynolds and D.F. Greboval, CIFA Technical paper 17, FAO 1988, ISBN 92-5-102742-0 (online version)
- M.L. Bianchini (1995). Species introductions in the aquatic environment: changes in biodiversity and economics of exploitation. Proc. World Fish. Congress (Athens, 1992), 3: 213-222.